Education: 

          B.A. - University of Washington (1966) 
          Ph.D - Tufts University (1973) 

Speciality:

Invertebrate neurobiology.

Research Interests:

Respiratory pumping in the marine snail, Aplysia.

Courses Taught:  

Cell Biology, Comparative Physiology, Physiological Ecology, Fish Physiology, Animal Behavior, Sociobiology of Reproduction, Comparative Animal Physiology, Evolutionary Biology, Biochemistry.